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The proton motive force can be expressed as where is the membrane potential and pH is the pH difference across the membrane, both expressed as the cytoplasmic minus the matrix values

doi: 10.1007/s12035-016-0170-2 134 TateWWalkerMSweetmanEHelliwellAPeppercornKEdgarCet al

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During the test phase, the preference for the novel arm was determined using a discrimination ratio (DR), which is calculated as the time spent exploring the novel arm divided by the total time spent in the start, familiar, and novel arms [novel/(novel + start+ familiar)]