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Hypoxia-induced HIF-1 activation can increase ROS levels through the elevation of NOX, or decrease ROS by inhibiting the tricarboxylic acid cycle [231]

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The results revealed that the levels of pro-inflammatory cytokines IL-6, IL-12, and TNF- secreted by cells treated with NP4 were 1.45, 5, and 1.78 times higher than those secreted by cells treated with NP2, respectively, further confirming the activation of macrophages