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Env Sci Technol 47:53275335 Johnson AAT, Kyriacou B, Callahan DL, Carruthers L, Stangoulis J, Lombi E, Tester M (2011) Constitutive overexpression of the OsNAS gene family reveals single gene strategies for effective iron and zinc biofortification of rice endosperm

What more-frequent treatment does do is degrade barrier function

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At physiological pH, these groups exist primarily in their deprotonated state
