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These three enzymes are involved in the processing of lignin and are shared by pathways involved in forming other key secondary metabolites, such as flavonoid, anthocyanin, and coumarin (Ali and McNear, GbMYBR1 , a trichomes repressor in Arabidopsis , led to reduced trichome density, reduced expressions of lignin-related genes such as PAL and 4CL , and decreased lignin content (Su et al., 2020)