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Radicals were not confirmed in living organisms until 1954

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23 Some of the proteins released from mitochondria into the cytoplasm include apoptosis-inducing factor and endonuclease G, both of which have nuclear localization signals and thus translocate to the nucleus, where the endonuclease cleaves DNA, resulting in DNA fragmentation

Notably, CH pQE1a- egtABCDE could produce 75 mg/L of ERG after 120 h of cultivation (Fig