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Polyphenols induce cell apoptosis by triggering reactive oxygen species (ROS) through two mechanisms 13 , direct formation of labile aroxil radical or a labile radox complex with a metal cation 42 and indirect activation of intracellular production of ROS by NADPH oxidases 43

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Interestingly, ToBRFV MP is less functional for viral cell-to-cell transport than TMV MP (Hak and Spiegelman 2021), suggesting that the possible evolution of ToBRFV via adaptation to Tm-2 2 tomato plants may have been associated with a fitness cost