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Generation of free radicals and oxidative stress Reactive oxygen species (ROS) are produced by living organisms due to normal cellular metabolism processes like cellular oxidation, cell regulation and signalling [4], and formed by the cells of aerobic organisms such as the electron transport chain, macrophages and peroxisomes [5]

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At elevated concentrations, H 2 O 2 operates in a dose-dependent manner to activate antioxidant defenses and to modulate the expression of stress-responsive genes, thereby exerting profound regulatory effects on plant growth and development ( Under copper stress, the activities of three key antioxidant enzymes in peanut seedling roots increased markedly: superoxide dismutase (SOD) rose by 11%, catalase (CAT) by 16%, and peroxidase (POD) by 75% ( Figure 1 )